Research Journal of Animal, Veterinary and Fishery Sciences __ _____________ ______ ____ __ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci. International Science Congress Association 1 Studies on Ichthyofaunal Diversity with special reference to Monthly and Seasonal variations of Fish Landings in glacial fed mountainous Goriganga River of Kumaun Himalaya, Uttarakhand, India Kumar Ashok Department of Zoology, Kumaun University Soban Sing h Jeena Campus Almora - 263601, INDIA Available online at: www.isca.in , www.isca.me Received 26 th February 201 4 , revised 6 th March 2014 , accepted 15 th April 201 4 Abstract Freshwater Ichthyofaunal diversity an d fish landings status of glacial fed Goriganga river, a major tributary of Kaliganga River of Pithoragarh district, Uttarakhand was studied from July 2006 to June 2008. Monthly sampling was carried out at three sampling stations (i.e. sampling station 1 - J auljibi, 600msl; Sampling station 2 - Baram 900msl and sampling station 3 - Madkot 1300msl). The results and observations of the present study reveals that apart from being glacial fed, Goriganga river harbours colourful ichthyo - fauna. Some 20 (Twenty) indige nous fish species belonging to twelve genera (Schizothorax, Tor, Barilius, Labeo, Punctius, Garra, Nemacheilus, Pseudecheneis, Glyptosternum, Homaloptera, Barbus and Mastacembelus) and five families (Cyprinidae, Botinae, Sisoridae, Homalopteridae and Masta cembelidae) were identified in the collection. The family Cyprinidae was dominant with 13(65%) fish species followed by Botinae with 3 (15%) species, Sisoridae with 2(10%) species and Homalopteridae and Mastacembelidae with 01(5%) species each. Monthly, se asonal and site wise variations in fish landings including similarity and dissimilarity index amog different fish species collected from selected sampling stations have also been discussed in the paper. Keywords: Ichthyofaunal , d iversity, f ish l andings, v ariations, g origanga, k umaun Himalaya. Introduction The Goriganga river originates from a dual source in a glacier near south of Untadhura ridge feeding the eastern branch and another glacier near Milam (3600 msl) just north east of Nanda Devi, feeding the western branch. This mixed glacier system offers breathe taking views that abounds in natural beauty. Goriganga is also fed by glaciers and streams flowing from the eastern slopes of the east wall of Nanda Devi sanctuary and those flowing west from th e high peaks of Panchachuli, Rajramba and Chaudhara, including Ralamgad and the Pyunsmi gadhera. The Kalabaland - Burfu Kalganga glacier also flows in to the Goriganga valley from the east. Goriganga captures the entire gradient of Himalayan habitats rangin g from subtropical sal to alpine meadows with in a distance of about 100 km of the river from its confluence to source. The breath taking beauty of Goriganga River, roaring down the zig - zag course, wide expenses of grassy meadows, a stupendous variety of f lora and fauna, above all, pure nature unsuilled, seem to beckon the beholder in to its folds in to a charmed world of virgin beauty. The total catchment area of the river is about 2230 sq.km. Out of which 346 sq.km is snow bound Joshi et.al. 1 and the whol e stretch of river is 100 km. and the Goriganga River runs through the entire length of the valley. Variety of fish species in any aquatic ecosystem is termed as ichthyo - diversity. Ichthyo - diversity is essential for stabilization of an aquatic ecosystem, protection of overall environmental quality and for understanding the intrinsic worth of all species in water as well as on the earth and in crude way, it is referred to listing of species, their number and status in a particular region. Millions of human beings suffer from hunger and malnutrition and fishes form a rich source of food and provide a meal to tide over the nutritional difficulties of man. In addition to serving as an important item of food, fishes provide several by - products to us. Fishes have formed an important item of human diet from time immemorial and are primarily caught for this purpose. Fish diet provides proteins, fat and vitamins A and D. A large amount of phosphorous and other elements are also present in it. They have a good taste a nd are easily digestible. However, it is an ephemeral phenomenon and is more related to the social and economic development of the region. In fact Fishes are such biotic components of the aquatic ecosystem which makes shuttle between aquatic ecosystem and socio economic sector of the people of the catchments. The fishery industry with its products and byproducts also gives a large scope for employment and socio - economic welfare of the society. From economic point of view, fish stands as the important source of protein food for the increasing population of world, especially for protein starved developing countries. Fishes are treasured resource both in terms of utility of food and as material for scientific study. Fishes occupy several different levels of the aquatic food chain, make up to over 40 % of the earth’s vertebrate species and are preyed upon by a few predatory animals and one another. Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 2 Leveque et.al. 2 have studied the global diversity of fish (Pisces) in fresh waters. According to them maximum 1403 5 fresh water fish species were reported from continent South America followed by 3553 species from Asia, 2945 species from Africa, 1411 species from North America, 330 species from Europe, 260 species from Oceania and minimum 206 fresh water fish species were reported from Russia. It is therefore, clear from the above observation made by Leveque et.al 2 . that continent Asia ranks second in fresh water fish diversity being South America ahead of it. Continent - wise fresh water fish diversity according to Leve que et.al 2 . shows the following distributional pattern as: South America (14035 species) � Asia (3553 species) �Africa (2945) � North America (1411) � Europe (330 species) � Oceania (260 species) � Russia (206 species). A considerable amount of literat ure is available on the fish and fishery of India by Jhingran 3,4 ; Mishra et.al. 5 ; Mishra and Sharma 6 ; Sharma et.al. 7 ; Anon. 8 ; and Sharma et.al. 9 . While Badola and Pant 10 reported 18 species of fishes from some districts of Uttaranchal, Badola 11 reported 4 3 species of fishes from some districts of Garhwal, Badola and Singh 12,13 reported 33 species of fishes from Chamoli districts of Uttaranchal, India. On the other hand Sing et.al. 14 have published a geographical and distributional list of ichthyofauna and reported a total of 68 species of fishes from Garhwal Himalaya. Das and Pathani 15 have also reported fish fauna of Kumaun lakes; Uniyal 16 reported 132 species of fishes from Uttaranchal State. While as per CAMP - 1998, 10 species are in the category of endan gered and need immediate protection and 10 species are exotic in nature and introduced in the state for various reasons. While 11 species are endemic to the Uttarakhand State, which act as a flag ship for balancing the aquatic ecosystem. Although a few rec ent literature on fish diversity have been consulted Pandey 17 , Thirupathaiah 18 , Gohil and Mankodi 19 , Kumar and Kumar 20 and sunita and Kanhere. 21 But there is a little information available on the fish and fisheries of the Goriganga river. Therefore, the pr esent venture is an attempt to study and to throw light on ichthyofaunal diversity, with monthly and seasonal variations of fish landings in the glacial fed mountainous Goriganga river of Kumaun Himalaya. Material and Methods Fishes were caught from different strata and mainly, the collections were made with the help of local fisher men and villagers using cast net, gill net, fatela jal, by hitting stones, angling using various types of bait viz. flour pellets, insects, earthworms etc. and various ty pes of local traps and Teep method (a very popular method in the local area used for capturing fishes after rainy season). The different types of fishing methods were used in different seasons. Fishes soon after collection were brought to the laboratories of Zoology department S. S. J. Campus Almora and preserved in 10% formalin solution in separate jars according to the size of the species. Small fishes were directly placed in the 10% formalin soltution, while large fishes were giving and incision in their abdomen and preserved. Meristic and morphometric characters were measured and identified up to the species level, with the help of standard keys and books by Day 22, Mishra 23 , Jayaram 24,25 , Qureshi and Qureshi 26 , Srivastava 27,28 , Talwar and Jhingran 29 an d Jhingran 30 . Monthly and seasonally, fish landing data were recorded from the local markets falling with in the vicinity of respective sites and the percentage of fish production was done on the basis of fish catch during the entire course of study. Res ults and Discussion Identification of fish: It is interesting to note that apart from being glacial fed, Goriganga river harbours colourful ichthyo - fauna. Some 20 (Twenty) indigenous fish species belonging to twelve genera (Schizothorax, Tor, Barilius, La beo, Punctius, Garra, Nemacheilus, Pseudecheneis, Glyptosternum, Homaloptera, Barbus and Mastacembelus ) and five families (Cyprinidae, Botinae, Sisoridae, Homalopteridae and Mastacembelidae) were identified in the collection from July - 2006 to June - 208 in t he Goriganga river t able - 1. Among these 20 fish species, four species of Schizothorax (S. richardsonii, S. plagiostomus, S. progastus and S. kumaonensis) exhibits 20.0 %; two species of Tor (Tort or and T. putitora) exhibited 10.0 %; two species of Baril ius (Barilius bendelisis and B. vagra) have constituted 10.0 %; one species of Barbus (Barbus chillinoides) constituted 5.0 %; two species of Labeo (Labeo dyocheilus and L. dero) constituted 10.0 %; one species of Punctius (Puctius ticto ) constituted 5.0 %; one species of Garra (Garra gotyla) constituted 5.0 %; three species of Nemacheilus (Nemacheilus rupicola, N. monatus and N. beavani) constituted 15.0 %; one species of Glyptosternum (Glyptosternum pectinopterum) constituted 5.0 %; one species of Pseu decheneis (Pseudecheneis sulcatus) constituted 5.0 %; one species of Homaloptera (Homaloptera brucei ) constituted 5.0 % and one species of Mastacembelus (Mastacembelus armatus) constituted 5.0 % in the present study. During first year (2006 - 07), the maxim um 20 fish species were recorded at spot - 1 Jauljibi followed by 15 fish species at spot - 2 Baram and the minimum 8 fish species were recorded at spot - 3 Madkot t able - 1; while during the second year (2007 - 08), the maximum 15 fish species were again recorded a t spot - 1 Jauljibi, 12 fish species at spot - 2 Baram and the minimum 8 species were again recorded at spot - 3 Madkot t able - 1. During both the years ichthyofaunal diversity of Goriganga river at three spots followed the distributional pattern as: spot - 1 (20 an d 15 fish species) � spot - 2 (15 and 12 fish species) � spot - 3 (8 and 8 fish species) during 2006 - 07 and 2007 - 08, respectively t able - 1. The spot - 1 (Jauljibi) appeared to be the richest site of ichthyo - faunal diversity (20 and 15 fish species) while spot - 3 Madkot appeared to be the poorest site of ichthyo - faunal diversity (08 and 08 fish species) during 2006 - 07 and 2007 - 08 respectively in the study t able - 1. Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 3 Table - 1 Qualitative composition of ichthyofauna at three spots in the Goriganga river during 2006 - 07 and 2007 - 08 July 2006 - June 2007 July 2007 - June 2008 S.No Genera / species Spot - 1. (Jauljibi) Spot - 2. (Baram) Spot - 3. (Madkot) Spot - 1. (Jauljibi) Spot - 2. (Baram) Spot - 3. (Madkot) 1. Schizothorax richardsonii + + + + + + 2. S. plagiostomus + + + + + + 3. S. progastus + + + + - - 4. S. kumaonensis + - - - - - 5. Tor tor + + - + - - 6. Tor putitora + + + + + + 7. Barrilius bendelisis + + + + + - 8. B. vagra + + - + + - 9. Labeo dero + + - - - - 10. L. dyocheilus. + - - + - - 11. Puntius ticto + - - - + - 12. Barbus chillinoides + + + + + + 13. Garra gotyla + + - + + + 14. Nemacheilus rupicola + + - + + + 15. N. monatus + - - + - - 16. N. beavani + + - - - - 17. Pseudecheneis sulcatus + + + + + + 18. Glypytosternum pectinopterum + + + + + + 19. Homaloptera brucei + - - - - - 20. Mastacembelus armatus + + - + + - Total 20 15 8 15 12 8 {+ = present and - = absent}. Table - 2 Seasonal distribution of ichthyofauna in the Goriganga river during 2006 - 07 and 2007 - 08 S.No Genera / speci es Summer Monsoon Winter 1. Schizothorax richardsonii +++ +++ +++ 2. S. plagiostomus +++ ++++ +++ 3. S. progastus ++ ++ ++ 4. S. kumaonensis ++ ++ ++ 5. Tor tor ++ ++ ++ 6. T. Putitora ++ ++ ++ 7. Barrilius bendelisis + + + 8. B. vagra + + + 9. Labeo dero ++ - - 10. L. dyocheilus. ++ - - 11. Puntius ticto + - - 12. Barbus chillinoides + - - 13. Garra gotyla ++ ++ ++ 14. Nemacheilus rupicola + + + 15. N. monatus + + + 16. N. beavani + + + 17. Pseudecheneis sulcatus * +++ * 18. Glyp ytosternum pectinopterun * +++ * 19. Homaloptera brucei - - + 20. Mastacembelus armatus + + + {+ = present; ++ = normal; +++ = abundance; - = absent and * = very rare} Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 4 Table - 3 Monthly variations in fish landings / production at three spots in the Gor iganga river during 2006 - 07 and 2007 - 08 Months Fish landing / Production (kg/month) July - 2006 to June - 07 July - 2007 to June - 2008 Jauljibi. (Spot - 1) Baram. (Spot - 2) Madkot. (Spot - 3) Jauljibi. (Spot - 1) Baram. (Spot - 2) Madkot. (Spot - 3) July 120.00 60.00 12.00 90.00 36.00 0.00 Aug 90.00 30.00 15.00 90.00 30.00 12.00 Sept 105.00 45.00 18.00 105.00 90.00 18.00 Oct. 90.00 30.00 24.00 90.00 60.00 15.00 Nov 90.00 60.00 24.00 90.00 51.00 30.00 Dec 90.00 60.00 15.00 90.00 30.00 12.00 Jan 90.00 45.00 27.00 105.00 45.00 15.00 Feb 75.00 30.00 15.00 120.00 75.00 24.00 Mar 108.00 60.00 24.00 75.00 30.00 18.00 Apr 90.00 60.00 24.00 96.00 30.00 09.00 May 120.00 75.00 30.00 126.00 87.00 30.00 Jun 135.00 60.00 12.00 75.00 30.00 0.00 Total 1203 615 240 1152 594 183 Annual Mean 100.25 51.25 20.00 96.00 49.5 15.25 Annual (%) 58.45% 29.88% 11.66% 59.72% 30.79% 9.48% S.D. 17.420 14.943 6.179 15.666 23.185 9.780 Table - 4 Seasonal variations in fish landing / production at three spots in the Goriganga river during 2006 - 07 and 2007 - 08 Seasons Fish landing / Production (kg/month) July - 2006 - June - 07 July - 2007 - June - 2008 Jauljibi (Spot - 1) Baram (Spot - 2) Madkot (Spot - 3) Jauljibi (Spot - 1) Baram (Spot - 2) Madkot (Spot - 3) Monsoon 112.5 48.75 14.25 90.0 46.5 7.5 Winter 90.0 48.75 22.5 93.75 46.5 18.0 Summer 123.25 56.25 23.25 104.25 55.5 20.25 Table - 5 Similarity and dissimilarity index among the different fish species identified in Goriganga river during 2006 - 07 and 2007 - 08 at all the spots in the study Genera Simila rity index Dissimilarity Index 2006 - 07 2007 - 08 2006 - 07 2007 - 08 1.Schizothorax 0.6 0.57 0.4 0.43 2. Tor 0.34 0.5 0.66 0.5 3. Barilius 0.4 0.0 0.6 1.0 4. Barbus 0.6 0.66 0.4 0.34 5. Labeo 0.6 0.0 0.4 1.0 6. Punctius 0.0 0.0 1.0 1.0 7. Garra 0.0 0.66 1.0 0.34 8.Nemacheilus 0.0 0.5 1.0 0.5 9. Pseudecheneis 0.6 0.66 0.4 0.34 10.Glyptosternum 0.6 0.66 0.4 0.34 11. Homapoptera 0.0 0.0 1.0 1.0 12. Mastacembelus 0.0 0.0 1.0 1.0 Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 5 Monthly qualitative composition of ichthyo - faunal diversity at different spots in the Goriganga river during 2006 - 07 and 2007 - 08 has been depicted in tables - 6,7,8 and figures 1 and 2. Figure - 1 Monthly variations in fish landings / production at three spots in the Goriganga river during 2006 - 07 Figure - 2 Monthly variations in fish landings / production at three spots in the Goriganga river during 2007 - 08 During first year (2006 - 07), the maximum (13) fish species were recorded in the month of July followed by December and September (12), January and April (l1), August and M ay (09), October, February , March and June (08) and the minimum (06) fish species were recorded in the month of July at spot - 1 Jauljibi, t able - 6 and f ig ure 1; at spot - 2 (Baram), the maximum (12) fish species were recorded in the months of December and Apr il followed by November and May (11), October (10), September and February (09), August, January and June (08), March (06) and the minimum (05) fish species were recorded in the month of July t able - 7 and f ig ure - 1, whereas at spot - 3 (Madkot), the maximum (0 6) fish species were recorded in the months of August and June followed by September, November and December (05), July, October, January and March (04) and the minimum (03) fish species were recorded in the month of May t able - 8 and f ig ure - 1. During second year (2007 - 08), the maximum (12) fish species were recorded in the month of October followed by September, November and December (10), August, January and May (09), June (08) and the minimum (07) fish species were recorded in the months of July, February, March and April at spot - 1 Jauljibi, t able - 6 and f ig ure - 2; at spot - 2 (Baram), the maximum (11) fish species were recorded in the month of April followed by November and December (10), January and May (08), July, October, February and March (07), September a nd June (06) and the minimum (05) fish species were recorded in the month of August t able - 7 and f ig ure - 2, while at spot - 3 (Madkot), themaximum (06) fish species were recorded in the months of July, October, April and June followed by September, November, D ecember March and May (05), January and February (04) and the minimum (03) fish species were recorded in the month of August t able - 8 and f ig ure - 2 in the present study. A maximum 13 (65.0 %) fish species belong to family Cyprinidae followed by 03 (15.0 %) species to Botinae, 02 (10.0 %) species to Sisoridae, and the minimum 01 (5.0 %) species to Homalopteridae and Mastacembelidae each were recorded in the present study t ables 6,7 and 8. On the basis of present survey it was observed that maximum number of f ish species were recorded from downstream as compared to the upstream or in other words, it was observed that ichthyofaunal - diversity increased from upstream to downstream (i,e. from high altitude to low altitude) in Goriganga river during 2006 - 07 and 2007 - 08 in the study t able - 1. Altitudinal distributions of Himalayan fishes are clearly distinguished by their body built modifications. The movement of fishes upwards and downwards from their natural habitats is affected by the water current (flood), temperat ure, nature of substrate, availability of food and their breeding activity. Extensive and intensive ichthyo - faunal surveys in Goriganga river depicted enormous potential of colourful ichthyo - fauna in its selected sampling stations and tributaries (Routes g ad / Gargia gad, Gossi gad, Shera gad etc.). The occurrence of fish species in different seasons of the year at all the selected spots is not same t able - 2. The fish species Schizothorax richardsonii, S. plagiostomus, and T. putitora were in abundance thro ughout the study period at all the selected spots, while cat fishes (Pseudecheneis sulcatus and Glyptosternum pectinopterum) were recorded more in numbers during rainy (monsoon) season than any other season of the year in the study t able - 2. Some species of fishes (Schizothorax richardsonii, S. plagiostomus, Tor putitora, Barbus chillinoides, Pseudecheneis sulcatus and Glyptosternum pectinopterum) were commonly present at all the selected spots t able - 2 during the entire course of study. Fish Landings: Month ly, seasonal and site - wise variations in fish landing have been recorded at three selected spots in the Goriganga river from June 2006 to July 2008 in the present study t ables - 3 and 4. The monthly fish landing (catch data) of two years (2006 - 07 and 2007 - 08) are furnished in the tabl - 3. The total fish landing Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 6 were recorded as 2058 kg (20.58 quintals) and 1929 kg (19.29 quintals) for 2006 - 07 and 2007 - 08, respectively in the Goriganga river. During first year (2006 - 07), monthly fish landing ranged from 75.0 kg (February) to 135.0 kg (June) at spot - 1, 30.0 kg (August, October and February) to 75.0 kg (May) at spot - 2 and from 12.0 kg (June and July) to 30.0 kg (May) at spot - 3; while during second year (2007 - 08) it ranged from 75.0 kg (March and June) to 120.0 k g (February) at spot - 1, 30.0 kg (August, December, March, April and June) to 90.0 kg (September) at spot - 2 and from 0.0 kg (June and July) kg to 30.0 kg (November) at spot - 3 in the Goriganga river Table - 3. The maximum fish landings during 2006 - 07 was reco rded at spot - 1 Jauljibi (1203 kg - 58.45%) followed by spot - 2 Baram (615 kg - 29.88%) and the minimum fish landings was recorded at spot - 3 (240 kg - 11.66%), while during second year (2007 - 08), the maximum fish landing was again recorded at spot - 1 Jauljibi (11 52 kg - 59.72 %) followed by spot - 2 Baram (594 kg - 30.78%) and the minimum (183 kg - 9.48 %) were again recorded at spot - 3 Madkot Table - 3. It was observed that, the maximum fish landing was recorded at spot - 1 (Jauljibi) and the minimum at spot - 3 (Madkot) dur ing 2006 - 07 and 2007 - 08 in the study and on the basis of this observation it is concluded that spot - 1 (Jauljibi) appeared to be the richest site of fish landing and spot - 3 (Madkot) appeared to be the poorest site of fish landing t able - 3. The highest annual mean values were recorded at spot - 1 Jauljibi (100.25 kg/month and 96.00 kg/month), while the lowest annual mean was recorded at spot - 3 Madkot (20.00 kg/month and 15.25 kg/month) during 2006 - 07 and 2007 - 08, respectively t able - 3. It is also observed that the fish landings decreased from downstream to upstream or altitudinaly fish landings increase with the decrease in altitude in the Goriganga river t able - 3. It was also observed that the fish landings was high during the summer season (123.25 kg/month, 56. 25 kg/month and 23.25 kg/month) and (104.25 kg/month, 55.5 kg/month and 20.25 kg/month) at spo - 1, spot - 2 and spot - 3 during first year (2006 - 07) and second year (2007 - 08), respectively t able - 4 and Figs - 3,4. While it was low in winter at spot - 1(90.00 kg/mont h) and in monsoon at spot - 2 (48.75 kg/month) and spot - 3 (14.25 kg/month) during 2006 - 07 t able - 4 and f ig ure - 3. The low fish production was recorded during monsoon season (90.00 kg/month, 46.5 kg/month and 7.5 kg/month) at spot - 1, spot - 2 and spot - 3, respecti vely during second year in the study t able - 4 and f ig ure - 4. The fish landing data showed fluctuations of fish catches in different months and seasons of the year, as well as year to year also t ables - 4. On the basis of present fish landing data, a slightly d ecreasing trend of fishery was observed in the Goriganga river which needs proper and immediate attention to save the colourful ichthyo - fauna of glacial fed mountainous Goriganga river of Kumaun Himalaya. Similarity and dissimilarity index among the diffe rent fish species identified at selected sampling stations in the Goriganga river during 2006 - 07 and 2007 - 08 has been attempted in the present study and depicted in the table - 5 and figs - 5 and 6. On the perusal of table - 5, it is clear that the maximum simil arity (s = 0.6) was observed among the species of Schizothorax, Barbus, Labeo, Pseudecheneis and Glyptothorax, while the minimum (s = 0.00) was among the species of Puctius, Garra, Nemacheilus, Homaloptera , and Mastacembelus during 206 - 07 f ig ure - 5. During 2007 - 08, the maximum similarity (s = 0.66) was among the species of Barbus, Garra, Pseudecheneis and Glyptothorax, while the minimum similarity (s = 0.00) was among the species of Barilius, Labeo, Homaloptera, and Mastacembelus f ig ure - 6 in the present s tudy. Figure - 3 Seasonal variations in fish landings / production at three spots in the Goriganga river during 2006 - 07 Figure - 4 Seasonal variations in fish landings / production at three spots in the Goriganga river during 2007 - 08 Discussion: The ob servations made on the ichthyofaunal - diversity and variations of fish landings of Goriganga river from July 2006 to June 2008, showed monthly, seasonally, yearly and altitudinal variations in the present study. During the course of study, some 20 indigenou s fish species which belong to coldwater species of fishes have been recorded from the different selected spots of Goriganga river, belongs to 12 genera ( Schizothorax, Tor, Barilius, Labeo, Punctius , Garra, Nemacheilus, Pseudecheneis, Glyptosternum, Homalo ptera, Barbus and Mastacembelus) and 05 families (Cyprinidae, Botinae, Sisoridae, Homalopteridae and Mastacembelidae. The Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 7 Cyprinidae family is richest (13 species - 65.00%) followed by Botinae (03 species - 15.00%), Sisoridae (02 species - 10.00%) and Homalopter idae and Mastacembelidae (01 species each - 5.00%). It is interesting to note that the Schizothorax richardsonii , S. plagiostomus, Tor putitora, Barbus chillinoides, Glypytosternum pectinopterum, Pseudecheneis sulcatus found and caught from all the spots but in different seasons of the year. Economically important fishes of the area are Schizothorax richardsonii , S. plagiostomus, Tor tor, T. putitora and Garra gotyla , the most relished fishes of the Kumaun Himalaya . It was observed that approximately 29 famil ies of fishermen from different villages of Goriganga basin (13 at Jauljibi, 9 at Baram and 7 at Bangapani and Sheraghat) and 04 families of fish farmers (one at Jauljibi, 02 at Gargia an 01 at Baram) are involved in fishing for their livelihood on thes e food fishes of the river. Figure - 5 Similarity and index among different species of fishes in the Goriganga river during 2006 - 07 Figure - 6 Similarity and index among different species of fishes in the Goriganga river during 2007 - 08 Various data on f ish landing have been recorded to access the monthly, seasonal and annual variations in their production at different selected sites from July, 2006 to June, 2008 in the Goriganga river. Total fish landings / production during both the years at all the sit es was recorded as 3987 kg (39.87 quintals). Fish landing / production during 2006 - 07 was 2058 kg (20.58 quintals - 51.61%) while it was 1929 kg (19.29 quintals - 48.25%) during 2007 - 08. A slight decreasing trend of fish landing / production was observed durin g second year in the study. Maximum fish landing / production (1203 kg – 30.17% and 1152 kg – 28.895) was recorded at Jauljibi (spot - 1), while minimum fish landing / production (240 kg – 6.01% and183 kg – 4.58%) was noticed at Madkot (spot - 3) during 2006 - 07 and 2007 - 08, respectively. Jauljibi appeared to be the richest site whereas Madkot to be the poorest site of fish production during both the years in the study. Spot wise variations of fish landing / production in Goriganga river from July 2006 to June 2008 followed the pattern as: spot - 1 (2355 kg - 59.06%) � spot - 2 (1209 kg - 30.32%) � spot (423 kg - 10.60%). Altitudinally, fish landing / production decreases with the increase in altitude (increases from upstream to downstream). Seasonally, maximum fish landing / production was observed in summer while minimum was in monsoon and winter, due to the reason that fish landing / production was greater during moderate water level (summer) than during increased water level (monsoon). Most of the fishes in Uttara khand state are potamodromous and few are migratory which are struggling hard to survive due to fragmentation of their aquatic habitats Sharma 31 . The fishery operations in Goriganga river are curtailed due to very fast water flow and the fisheries include only the fish species with either strong power of locomotion or special organs of attachment. Therefore, snow trout, Mahseer and some minor carps constitute the major fishery of the Goriganga river. Fish species belonging to Schizothorax genera makes the m aximum contribution to the fishery of the concerned water body at all the spots during both years (2006 - 07 & 2007 - 08). It is well known that Schizothorax richardsonii is highly esteemed as food fish in most of the Himalayan states and highly preferred by f ish eaters. The species of Labeo, Pseudecheneis, Barbus and Glyptosternum, also forms small part of the fish community of the Goriganga river. The species of Noemacheilus, Puntius, Barilius, Homaloptera and Mastacembelus are rare in the main river, but th ey occur in abundance in its tributaries such as Routes gad, Gossi gad, Shera gad etc. The glacial fed Goriganga river flow with high speed among boulders between the hard rocks at high altitude. The rocky bottom which results in the occurrence of rare f ishes hiding under crevices and boulders. The siluroid fishes (cat fishes - Pseudecheneis and Glyptosternum ) are found in these conditions and they have developed special adhesive organs for attaching to any hard surface, so these fishes hide themselves bel ow boulders with the help of such specialized organs. It was observed that the catch percentage of such fishes were high during the rainy season. The reason behind this is that during rainy season due to furious flow of the river the fishes are detached fr om the boulders and come to the surface, so caught easily. After rainy season the flow becomes moderate and the Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 8 fishes again inhibits the crevices of rocky bottom and becomes difficult to catch these siluroid fishes. In the winter months when the water lev el comes down hammering method, hand pinking, diversion of rivulets water are used to catch fishes from the river and its tributaries. Table - 6 Monthly qualitative composition of ichthyo - faunal diversity at spot - 1 (Jauljibi) in the Groriganga river duri ng 2006 - 07 and 2007 - 08 Monthly qualitative composition of ichthyo - faunal diversity at spot - 1 (Jauljibi) in the Groriganga river during 2006 - 07 and 2007 - 08. July Aug Sept Oct Nov Dec Jan Feb March Apr May Jun Cyprinidae: I II I II I II I I I I II I II I II I II I II I II I II I I I Schizothorax richardsonii + + + + + + + + + + + + + + + + + + + + + + + + S. plagiostomus + + + + + + + + + + + + + + + + + + + + + + + + S. progastus - - - - - - - - + - + - + - - + + - - - - - - - S. kumaonensis - - - - - - - - + - - - - - - - - - - - - - - Tor tor - + + + + - + - + + - + + + - - - - + - + - - - Tor putitora + + + + + + + + + + + + + + + + + + + + + + + + Barrilius bendelisis - - + + + + + + + + + + + + + + - + + - - - - - B. vagra - - - - + - - + - + + - - - - - - - + + + + - - Labeo dero - - + + + + + - - - - - + - + - - - + - - + - + Labeo dyocheilus - - - - + - - + - - + - - + - - - - - + - + + + Puntius ticto - - - - - - - - - - - + + + - - - - + + + + - - Barbus chillinoides + + - - + + - + + - + + - + + + - + + - + - + + Garra gotyla + - + + + + + + + + + + + - + - + + + - + - + - Botinae: Nemacheilus rupicola - - - + + + + + + + + + + - - + + + - + - + - - N. monatus - - - - - - - + - + - + - - - - - - - - - - - - N. beavani - - - - - + - + + - + - - - + - + - - - - - - - Sisoridae: Pseudecheneis sulcatus + + + - - - - - - - - - - - - - - - - - - - + + Glypytosternum pectinopterun + + + - - - - - - - - - - - - - - - - - - - + + Homa popteridae: Homaloptera brucei - - - - - - - - + - - - - - - - - - - - - - - - Mastacembelidea: Mastacembelus armatus - - - + + + - + + + + + + + - - + - + - + + - - Total 07 07 09 09 12 10 08 12 13 10 12 10 11 09 08 07 08 0 7 11 07 09 09 08 08 (I = during 2006 - 07; II = during 2007 - 08; + = present and - = absent) Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 9 Table - 7 Monthly qualitative composition of ichthyo - faunal diversity at spot - 2 (Baram) in the Groriganga river during 2006 - 07 and 2007 - 08 Genera / species Monthly qualitative composition of ichthyo - faunal diversity at spot - 2 (Baram) in the Groriganga river during 2006 - 07 and 2007 - 08 July Aug Sept Oct Nov Dec Jan Feb March Apr May Jun Cyprinidae: I II I II I II I II I II I II I II I II I II I II I II I II Schizothorax richardsonii + + + + + + + + + + + + + + + + + + + + + + + + S. plagiostomus + + + + + + + + + + + + + + + + + + + + + + + + S. progastus - - + - + - + + + + + + - + + - + - + - - - + - S. kumaonensis - - - - - - - - - - - - - - - - - - - - - - - - Tor tor - - + - - - - - - - + - + - + - + + + - + + - - Tor putitora + - + + + + + + + + + + + + + + - + + + + + + + Barrilius bendelisis - - - - + - + - + + + + - - + - + + + + + - - - B. vagra - - - - + - - - + - - + + + - - - + - + - + - - Labeo dero - - - - - - + - + - + - - - - - - - + - + - - - L. dyocheilus - + - - - - - - - + - + - + - - - - - + - + - - Puntius ticto - - - - - - - - - - - - - - - - - - - - - - - - Babbus chillinoides - + - + + + + + + + + + + - + + - - + + + + + - Garra gotyla - + + + - + + + + + + - + + + + + + + + + + + - Botinae: Nemacheilus rupicola - - - - + - + - + + + + - - + + - - + + + - - + N. monatus - - - - - + - + - + - - - + - - - - - + - - - - N. beavani - - - - - - + - - - + - + - - - - - + - + - - - Sisoridae: Pseudecheneis sulcatus + + + - - - - - - - - - - - - - - - - - - - + + Glypytosternum pectinopterun + + + - - - - - - - - - - - - - - - - - - - + + Homapopterida e: Homaloptera brucei - - - - - - - - - - - - - - - - - - - - - - - - Mastacembelidea: Mastacembelus armatus - - - - + - - - + - + + - - - + - - + + + - - - Total 05 07 08 05 09 06 10 07 11 10 12 10 08 08 09 07 06 07 12 11 11 08 08 06 (I = during 2006 - 07; II = during 2007 - 08; + = present and - = absent) According to Dwivedi et.al. 32 “In inland fisheries of India and future prospectus”, they reported that at global level 85 % of the world fish production comes from the ocean and only 15% is contrib uted by fresh water. However, in India the picture is different. During 1950s marine fisheries contributed about 70 % and inland fisheries only 30 %. Now after over 50 years, inspite of low investments in inland fisheries sector, inland fish production has grown at a faster rate as compared to marine fisheries. During the year 2003, it was estimated that the contribution of fish production from inland waters had increased from 30 % to 50 % and contribution of marine fish production to total Indian fish prod uction has come down from 70 % to 50 %. It is estimated that in next ten years inland fisheries of India will contribute 60 % of total Indian fish landings. Further it may be noted that the average selling price of fresh water fishes varies between Rs. 4 0 to Rs. 90 per kg., whereas marine fishes are sold at around Rs. 30 to Rs. 50 per kg, indicating that fresh water fishes are priced high as compared to marine fishes. Few years ago ICAR had made a comparative study of income from different sectors, which showed that agriculture, dairy and fishery were most productive, providing a return of the order of Rs. 72,000 per ha. Dwivedi et.al. 33 It seems that glacial fed Goriganga river is suitable for fish diversity and its population even at the present time but the size of large fishes is dwindling in the river. it may have many causes (pollution, dynamiting, mass fish killing, low meshed nets, poisoning, fish catch during breeding season, water diversion, use of agricultural biocides, unawareness to fisherme n etc.). In Kumaun Himalaya, development of fisheries have not been taken up seriously by the concerning authorities. A large number of derelict water bodies are untouched and are in the verge of swampification. These water bodies require urgent attention for demonstration and extension of scientific fish culture techniques to make people aware towards the fish farming to enhance fish production Pathani 33 . Goriganga river harbours rich fish fauna at present and if the present status of the river system is m aintained through strategic water quality management measures (because fishes live in intimate contact with their environment and are therefore, very susceptible to physical and chemical changes) , it will ensure a sustainable aquatic environment capable of producing the fish protein food of the populace and enhance the people’s livelihood. Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 10 Table - 8 Monthly qualitative composition of Ichthyo - faunal diversity at spot - 3 (Madkot) in the Groriganga river during 2006 - 07 and 2007 - 08 Genera / species Monthly quali tative composition of Ichthyo - faunal diversity at spot - 3 (Madkot) in the Groriganga river during 2006 - 07 and 2007 - 08. July Aug Sept Oct Nov Dec Jan Feb March Apr May Jun Cyprinidae: I II I II I II I II I II I II I II I II I II I II I II I II Schizothorax richardsonii + + + + + + + + + + + + + + + + + + + + + + + + S. plagiostomus + + + + + + + + + + + + + + + + + + + + + + + + S. progastus - - - - + - + - - - + - - - + - - - - - - - - - S. kumaonensis - - - - - - - - - - - - - - - - - - - - - - - - Tor tor - - - - - - - - - - - - - - - - - - - - - - - - Tor putitora - + + - + + + + + - + - + + - + + + - + + + + - Barrilius bendelisis - - - - - - - - + - + - + - - - - - + - - - - - B. vagra - - - - - - - - - - - - - - - - - - - - - - - - Labeo dero - - - - - - - - - - - - - - - - - - - - - - - - L. dyocheilus. - - - - - - - - - - - - - - - - - - - - - - - - Puntius ticto - - - - - - - - - - - - - - - - - - - - - - - - Barbus chillinoides - - + - + + - + + + - + - + - + + + - + - + + - Garra gotyla - + - + - - - + - + - + - - - - - + - + - + - + Botinae: Nemacheilus rupicola - - - - - + - + - + - + - - - - - - - + - - - - N. monatus - - - - - - - - - - - - - - - - - - - - - - - - N. beavani - - - - - - - - - - - - - - - - - - - - - - - - Sisoridae: Pseudecheneis sulcatus + + + - - - - - - - - - - - - - - - - - - - + + Glypytosternum pectinopterun + + + - - - - - - - - - - - - - - - - - - - + + Homapopter idae: Homaloptera brucei - - - - - - - - - - - - - - - - - - - - - - - - Mastacembelidea: Mastacembelus armatus - - - - - - - - - - - - - - - - - - - - - - - - Total. 04 06 06 03 05 05 04 06 05 05 05 05 04 04 03 04 04 05 03 06 03 05 06 05 (I = during 2006 - 07; II = during 2007 - 08; + = present and - = absent) For conservation of dwindling fish diversity, ecological as well as anthropogenic stresses must be removed at t he first phase, scientific gear and mesh sizes needs to be regulated to protect the juvenile fishes. Mass awareness for conservation of valuable fish stocks may be explored through extension and by voluntary agencies. Further the involvement of entire scie ntific community in the drive is quite necessary and the involvement of voluntary organizations would always remain highly significant in accomplishing the task. Causes of decline of fish and fishery in Kumaun rivers with various suggestions have already b een made by Pathani 34,35 . Various causes like unscientific methods of fish catching, unscientific human interferences, over fishing, dynamiting, fishing during spawning season, use of bleaching powder and ichthyotoxic plants in the shallow breeding groun ds of tributaries were assessed to be responsible for declining trend of fishery in the Goriganga river. These are the cruel methods by which not only the adults of edible size are killed, but all the stages of fish life are destroyed and the natural habit at of the fish is also polluted. The pollution of the river by flash floods, landslides, and soil erosion etc. have also been responsible for the depletion of the fish fauna Badola and Singh 36 . Habitat destruction, alteration in ecological conditions of aq uatic resources, deforestation, soil erosion, silting, water currents alterations, forest denudation, unscientific developmental activities and over - grazing along slopy catchments erode soil structure and cause decline in fish production. Siltation from ca tchment areas of the river has destructed breeding and feeding grounds of many fishes. Pesticides washing from agricultural fields have also been created detrimental environment for fish life in the water. The fragile mountain ecosystems and native fish g ermplasm resources are our national wealth and pride. All flora and fauna of the upland water system have coevolved during the course of Research Journal of Animal, Veterinary and Fishery Sciences ___ _ _ _______________________________ ____ ISSN 2320 – 6535 Vol. 2 ( 4 ), 1 - 1 2 , April (201 4 ) Res. J. Animal, Veterinary and Fishery Sci . International Science Congress Association 11 organic evolution for their mutual benefit. Any species getting extinct would upset the ecological balance causing detr imental to each and every other species. Let us all join in protection and conservation of fish diversity in particular and biodiversity in general not only in Goriganga river but in the country as a whole, because it is the need of the hour. On the behalf of the present study, it was observed that for proper fishery management, it is desirable to impose a ban on illegal fishing and fishing during the breeding season (pre - spawning and spawning season) should be strongly prohibited and the most important thi ng is that provisions should be made by the concerned authorities or department to educate the people especially the fishermen, fish farmers and fish vendors of the catchment area (Goriganga basin) for the conservation of existing colourful ichthyofana of the Goriganga river. Conclusion The present study would give preliminary knowledge on the Ichthyofaunal diversity and reasons for monthly, seasonal and site wise variations of fish landings at different sampling stations. In the present investigation the qualitative composition of fish species during July - 2006 to June - 2008, it was cyprinidae 13 species 65 %, Botinae 3 species 15%, Sisoridae2 species 10% and Homalopteridae and Mastacembelidae with 01species each 5%. The fish species Schizothorax richardson ii, S. plagiostomus, and T. putitora were in abundance throughout the study period at all the selected spots, while cat fishes (Pseudecheneis sulcatus and Glyptosternum pectinopterum) were recorded more in numbers during rainy (monsoon) season than any oth er season of the year in the study. Some species of fishes (Schizothorax richardsonii, S. plagiostomus, Tor putitora, Barbus chillinoides, Pseudecheneis sulcatus and Glyptosternum pectinopterum) were commonly present at all the selected spots during the en tire course of study. It was also observed that ichthyofaunal - diversity including quantitative estimation increased with the decrease in altitude in the river. 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